In a story on the BBC website, about the streaked tenrecs of Madagascar, I find the following:
Unique hedgehog-like mammals have been filmed using their quills to communicate.
I wonder if they’ll eventually progress to fountain pens, biros, and on to other, more modern, technology.
(Yes, the photo includes all sorts of irrelevant stationery items, but it was the only one I had in my files that featured a quill pen. I think it may be a porcupine quill.)